Long-horned, 'humpless' domestic cattle were well established in the Nile Valley by 4000 BC. These cattle, known as the Egyptian or Hamitic Longhorn, appear in pictographs in Egyptian pyramids. Over the next 2000 years, the Egyptian Longhorn migrated with its owners from the Nile to Ethiopia, and then down to the southern reaches of Africa.
When these Zebu reached the region now known as Ethiopia and Somalia, they were interbred with the Egyptian Longhorn. This combination produced the Sanga which spread to the Sudan, Uganda, Kenya, and other parts of eastern Africa, becoming the base stock of many of the indigenous African breeds. The Sanga demonstrated most of the typical Zebu characteristics, such as pendulous dewlap and sheath, upturned horns, and a neck hump of variable size. Modern descendants of the Sanga, however, vary greatly in size, conformation, and horns, due to differing selection pressures by different tribes.
Particularly remarkable are the cattle found in Uganda, Rwanda, and Burundi. In Uganda, the Nkole tribe's Sanga variety is known as the Ankole. In Rwanda and Burundi, the Tutsi tribe's Sanga variety is called the Watusi. The Rwanda common strain of Watusi is called Inkuku. The giant-horned strain, owned by the Tutsi kings and chiefs, is called the Inyambo, though some current tribal reports claim that this type is now extinct.
Traditionally, Ankole-Watusi were considered sacred. They supplied milk to the owners, but were only rarely used for meat production, since an owner's wealth was counted in live animals. Under traditional management, the Ankole cow was grazed all day, then brought home to her young calf. The calf was allowed to suckle briefly to stimulate milk letdown, then the cow was milked by the herdsman. The calf suckled after hand-milking was finished and was again separated from its mother. The process was repeated in the morning. This minimal nourishment of calves resulted in high death rates in the young. Milk production was not high, with a typical cow producing only 2 pints of milk daily, although an exceptional one could manage up to 8 pints. In addition, the lactation period was short. Over the last 10 years, the national government has attempted to select for animals which produce more milk and have better meat production. Famine and disease, as well as the conflict with traditional practices, have slowed this effort.